I am an economist focusing on the long-run evolution of populations, through factors such as diversity, fertility,
and migration, and their relationship to economic growth and development. My work explores the deeply rooted historical
and demographic drivers behind modern economic and social outcomes, including the influence of ancient human migration
(Out of Africa) on institutional and technological variation, the trade-offs between child quantity and quality,
and the role of natural resources in local development. By examining genealogies, historical data,
and contemporary measures of diversity, I aim to shed light on how past events and evolutionary forces shape
our present economic conditions and future prospects.
PhD in Economics, University of Copenhagen (2013)
Associate Professor, University of Copenhagen
Visiting Scholar, Brown University
Research Affiliate, Centre for Economic Policy Research (CEPR)
My research explores interconnected themes in long-term economic development, demographics, and health. Below are the major themes that characterize my work.
Population Diversity and Long-Run Development
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How ancient migration patterns shape modern economic, political, and social outcomes.
This research theme explores how population characteristics shaped by ancient human migration patterns influence contemporary economic, political, and social outcomes. My work establishes a fundamental link between the genetic diversity resulting from human migration out of Africa tens of thousands of years ago and modern phenomena such as civil conflict, institutional development, and economic performance.
Using novel datasets on population diversity and advanced econometric methods, I've shown that diversity has both beneficial and detrimental effects on societies. While intermediate levels of diversity foster innovation and creativity through the interplay of varied cognitive approaches, excessive diversity can lead to reduced social cohesion, interpersonal trust, and increased civil conflict. These insights help explain persistent differences in the wealth of nations and provide a deeper understanding of societal development.
My research in this area contributes to our understanding of how deeply-rooted factors continue to influence modern economic and political outcomes, with important implications for policy approaches to managing diversity in contemporary societies.
How fertility decisions influence economic development through quantity-quality tradeoffs.
This research theme investigates the relationship between demographic behaviors and economic outcomes, with particular focus on the quantity-quality tradeoff in children. My work uses unique historical datasets, including reconstructed genealogies spanning centuries, to demonstrate how reproductive decisions influenced human capital formation and contributed to modern economic growth.
A key finding from this research is that moderate fecundity maximized long-run reproductive success. Using data from Quebec covering 1608-1800, I've shown that while higher fecundity led to more children in the short term, families with moderate fecundity achieved greater long-run reproductive success. This suggests natural selection favored moderate fertility rates, supporting a quantity-quality tradeoff in which parents invested more resources in fewer children.
I've also uncovered evidence that pre-industrial families actively controlled their fertility through birth spacing, challenging traditional Malthusian narratives. This work provides important insights into the demographic transition and demonstrates that even historical populations employed sophisticated fertility control strategies in response to economic conditions.
How historical shocks, resources, and market conditions shape long-term development.
This research theme examines how historical shocks, resource access, and economic conditions have influenced long-term development paths. My work connects early-life conditions, resource availability, and market dynamics to explain economic transitions and persistent effects that continue to shape modern economies.
Using empirical evidence from historical England and Scandinavia, I've demonstrated that regions experienced a "post-Malthusian" phase where population and income grew simultaneously despite diminishing returns. This research supports transitional growth theories that bridge Malthusian stagnation and modern economic growth, providing insights into the mechanisms driving the Industrial Revolution.
My work on natural resources has challenged traditional "resource curse" narratives by showing that localities closer to oil fields in Brazil enjoy higher per capita income, independent of oil royalties. This suggests "indirect linkage" effects where resource proximity creates positive economic spillovers, informing our understanding of how natural resources can contribute to local economic development.
Investigating the interplay between health factors, biology, and socioeconomic outcomes.
This research theme focuses on the intersection of health, biology, and economic outcomes. My interdisciplinary work spans from historical health shocks to contemporary medical studies, investigating how biological factors influence both short and long-term wellbeing across different populations.
Through collaboration with medical researchers, I've contributed to understanding how antihypertensive medication affects glaucoma onset, how brain responses correlate with intelligence, and the fertility patterns of specific populations. This research uses advanced statistical methods applied to large-scale medical and population datasets to identify causal relationships in health outcomes.
Historical health research is also a key component of this theme. My work on the English famine of the 1720s demonstrated that early-life adversity caused permanent health damage, with affected individuals experiencing higher mortality risks into adulthood. These findings contribute to our understanding of the lasting impact of health shocks and have implications for how societies should respond to health crises today.
Inferring Glaucoma Status from Prescriptions, Diagnoses, and Operations Data: A Danish Nationwide Study
PLOS ONE, 2023 | with A. Horwitz, J. Rovelt, H. Horwitz, C. Torp-Pedersen, M. Kolko
Summary: This study analyzes prescription, diagnosis, and operation data to accurately infer glaucoma status in a large Danish population dataset, providing a valuable method for identifying glaucoma cases in medical research.
The Evolutionary Origins of the Wealth of Nations
Handbook of Historical Economics (North Holland), 2020 | with Quamrul Ashraf and Oded Galor
Summary: This chapter explores how evolutionary processes and population dynamics have influenced the long-term economic development of nations, examining factors such as genetic diversity, geographical conditions, and historical migration patterns.
Human Genealogy Reveals a Selective Advantage to Moderate Fecundity
Nature Ecology & Evolution, 3(5), 853–857, 2019 | with O Galor
Summary: Using a reconstructed genealogy of nearly half a million individuals in Quebec (1608–1800), we show that while higher fecundity led to more children, moderate fecundity maximized long-run descendants. Evolutionary forces decreased fecundity over time, suggesting an advantage for having children at an intermediate rate. Mechanisms include improved child quality and marriageability.
Galor, O., & Klemp, M. (2019). Human Genealogy Reveals a Selective Advantage to Moderate Fecundity. Nature Ecology & Evolution, 3(5), 853-857. https://doi.org/10.1038/s41559-019-0846-x
@article{galor2019human,
title={Human Genealogy Reveals a Selective Advantage to Moderate Fecundity},
author={Galor, Oded and Klemp, Marc},
journal={Nature Ecology \& Evolution},
volume={3},
number={5},
pages={853--857},
year={2019},
publisher={Nature Publishing Group},
doi={10.1038/s41559-019-0846-x}
}
Post-Malthusian Dynamics in Pre-Industrial Scandinavia
Scandinavian Journal of Economics, 118(4), 841-867, 2016 | with NF Møller
Summary: Similar to England, Denmark, Norway, and Sweden displayed a post-Malthusian phase where incomes and population grew despite diminishing returns, lending broader support to transitional post-Malthusian growth theory.
Picking Winners? The Effect of Birth Order and Migration on Parental Human Capital Investments in Pre-Modern England
European Review of Economic History, 17(2), 210–232, 2013 | with C Minns, P Wallis, and J Weisdorf
Summary: Linking apprenticeship records with family reconstitution data, we find that eldest sons of non-gentry were often apprenticed, implying parental choice by aptitude rather than inheritance norms. Return migration was frequent, suggesting parents anticipated direct benefits from children's skill acquisition.
The Lasting Damage to Mortality of Early-Life Adversity: Evidence from the English Famine of the Late 1720s
European Review of Economic History, 16(3), 233–246, 2012 ("Editor's Choice") | with J Weisdorf
Summary: Individuals born during the severe 1720s English famine suffered higher mortality risks well into adulthood, suggesting a permanent negative health shock. Those born just before or after did not share this fate.
Prices, Wages, and Fertility in Pre-Industrial England
Cliometrica, 6(1), 63–77, 2012
Summary: This study explores the relationship between real wages, prices, and fertility in pre-industrial England. Using cointegration and error-correction modeling, it finds that fertility responded negatively to increases in agricultural prices and positively to increases in real wages, supporting Malthusian theory.
Human Colonization of Earth and Diverse Societal Developments (in Danish)
Danish Journal of Economics, 2022
Summary: This research provides an overview of how ancient migration from Africa
60–90 thousand years ago shaped global population diversity, influencing modern economic performance,
civil conflicts, political institutions, and interpersonal trust.
Historical Instruments and Contemporary Endogenous Regressors
Journal of Development Economics, 149, 2021 | with G Casey
Summary: We develop an augmented IV estimator to address time gaps between historical instruments
and contemporary regressors, applying it to institutions and Protestantism. Our findings suggest
somewhat smaller, but still important, long-run effects than standard estimates reveal.
The Journal of Clinical Endocrinology & Metabolism, 106, 442–449, 2020 | with J Windfeld-Mathiasen, K. Dalhoff, J. Andersen, A. Horwitz, H. Horwitz
Summary: Investigating 545 men testing positive for anabolic steroids in Denmark, we find a
26% lower fertility rate prior to doping. However, fertility partially recovers, suggesting AS-induced
infertility may be reversible. Usage of assisted reproduction and ultimate parenthood rates are
comparable to controls.
Econometrica, 88(2), 727–797, 2020 | with E Arbatlı, Q Ashraf, and O Galor
Summary: This research establishes that interpersonal population diversity has been pivotal to
the emergence and severity of intrasocietal conflict, measured by an exogenous variation from ancient human
migration out of Africa. Mechanisms include mistrust, divergent preferences, and reduced cohesion.
Arbatlı, C. E., Ashraf, Q. H., Galor, O., & Klemp, M. (2020). Diversity and Conflict. Econometrica, 88(2), 727-797. https://doi.org/10.3982/ECTA13734
@article{arbatli2020diversity,
title={Diversity and Conflict},
author={Arbatl{\i}, Cemal Eren and Ashraf, Quamrul H and Galor, Oded and Klemp, Marc},
journal={Econometrica},
volume={88},
number={2},
pages={727--797},
year={2020},
publisher={Wiley Online Library},
doi={10.3982/ECTA13734}
}
European Economic Review, 127, 2020 | with M Gradstein
Summary: Using Brazilian data on oil reserves, we find that localities closer to oil fields
enjoy higher per capita income. This positive impact is independent of oil royalties, consistent with
"indirect linkage" effects rather than a straightforward resource curse.
Fecundity, Fertility and the Formation of Human Capital
Economic Journal, 129, 925–960, 2019 | with J Weisdorf
Summary: Exploiting genealogical data for 16th–19th century England, we measure parental
reproductive capacity via the interval between marriage and first birth. While higher fecundity yielded
larger sibling sets, it reduced children's socio-economic outcomes, consistent with a quantity–quality
trade-off at the origin of modern economic growth.
Klemp, M., & Weisdorf, J. (2019). Fecundity, Fertility and the Formation of Human Capital. The Economic Journal, 129(618), 925-960. https://doi.org/10.1111/ecoj.12589
@article{klemp2019fecundity,
title={Fecundity, Fertility and the Formation of Human Capital},
author={Klemp, Marc and Weisdorf, Jacob},
journal={The Economic Journal},
volume={129},
number={618},
pages={925--960},
year={2019},
publisher={Oxford University Press},
doi={10.1111/ecoj.12589}
}
Brain Responses to Passive Sensory Stimulation Correlate With Intelligence
Frontiers in Aging Neuroscience, 11, 2019 | with A. Horwitz, H. Horwitz, M. D. Thomsen, E. Rostrup, E. L. Mortensen, M. Osler, M. Lauritzen, K. Benedek
Summary: Using EEG-based steady-state responses (SSRs) to passive audiovisual stimulation in
middle-aged men, we find alpha–gamma power differences strongly correlate with intelligence. This
link is robust to confounders and consistent with earlier-life IQ measurements.
Further Evidence of Within-Marriage Fertility Control in Pre-Transitional England
Demography, 56(4), 1557–1572, 2019 | with F Cinnirella and J Weisdorf
Summary: Building on prior evidence for parity-dependent birth spacing, we find further indications of within-marriage
fertility control in 17th–19th century England. This research adds to the understanding of pre-transitional
demographic behaviors preceding the demographic revolution.
Malthus in the Bedroom: Birth Spacing as a Preventive Check Mechanism in Pre-Modern England
Demography, 54(2), 413–436, 2017 | with F Cinnirella and J Weisdorf
Summary: Using family reconstitution data for England (1540-1850), we show that worsening economic conditions were associated with increased birth spacing. This relationship is stronger after 1750, and concentrated among poorer families, suggesting deliberate fertility control was a preventive check used by families to manage economic stress.
Antihypertensive Medication Postpones the Onset of Glaucoma
Hypertension, 69(2), 202–210, 2017 | with A. Horwitz, J. Jeppesen, J. Tsai, C. Torp-Pedersen, M. Kolko
Summary: Analyzing 2.6 million Danes reveals that while hypertension is associated with
higher glaucoma risk, antihypertensive medication reduces that risk in a causal sense. This supports
a protective effect of blood pressure treatment on glaucoma onset.
Visual Steady State in Relation to Age and Cognitive Function
PLOS ONE, 12(2), 2017 | with A. Horwitz, M. Thomsen, I. Wiegand, H. Horwitz, M. Nikolic, L. Rask, M. Lauritzen, K. Benedek
Summary: We observe alpha–gamma band differences in visual steady-state potentials in older adults, strongly correlating with intelligence. The correlation persists when controlling for speed of processing and other factors, indicating a potential clinical tool for cognitive assessment.
with O Galor | A more recent (2025) version is available on request
Summary: Proposes that population diversity, shaped by the exodus of Homo sapiens from Africa,
contributed to autocratic institutions historically. Diversity can enhance productivity but also fosters
scope for domination, with lasting implications for modern political systems.
with Q Ashraf and O Galor | A more recent (2025) version is available on request
Summary: We document a hump-shaped relationship between within-group genetic diversity and
productivity, capturing both the innovative benefits and frictions that diversity can produce. Cross-ethnic
data at subnational levels confirm the trade-off across economic outcomes.
ESTETA - Stata Command for Estimating Long-Run Effects Using Historical Instruments
2021 | with Gregory Casey
Description: This Stata command implements the augmented IV estimator developed in our JDE paper "Historical Instruments and Contemporary Endogenous Regressors." It addresses limitations of conventional IV regression when measuring persistent effects with historical instruments by accounting for the persistence of endogenous factors over time.
MATLAB Program for Numerically Solving Overlapping Generations (OLG) Models
2009 | with Christian Groth
Description: This program visualizes the phase space of Overlapping Generations (OLG) models, including scenarios with multiple
intertemporal equilibria. It does not compute full transition paths, but highlights equilibria and dynamic behaviors.